Photo 1. 1Y Goldfinch showing a few unmoulted juvenile feathers in front of the eye. |
Photo 2. 1Y Goldfinch showing 1 unmoulted juvenile feather below the eye and a few others still in pin. |
Photo 3. Known adult Goldfinch with 4 pure white feathers below and behind the eye (unusual). |
When I started to photograph every Goldfinch that I caught I didn’t expect to learn as much as I have. One example being that the moult of both adults and juveniles can extend well into December with the feathers of the face being the last to be replaced. This has particular implications for sexing birds but that will be the subject of another post if I ever get round to writing it. This post has been difficult to put together and has already had several false starts. What photos do you use when you have so many to choose from and what level of knowledge do you pitch it at ? I am likely to come back to this post from time to time to add photos or statistics so it should be seen as a work in progress. Hopefully some of you, especially other ringers, may find it of interest.
Most juvenile Goldfinches in the UK only undergo a partial post juvenile moult but it is thought that a very small proportion may have a complete moult like adults. The proportion of juveniles that undergo a complete moult isn’t easy to measure but is likely to be insignificant in the UK, if it happens at all. I didn’t encounter any juveniles in active primary moult and only caught 3 (<1%) that had replaced a few central primaries. However I did find that many replaced one or more tertials and some of their tail feathers.
Photo 4. 1Y Goldfinch with all new tertials, the longest tertial is not fully grown. The new glossy black tertails contrasting with the juvenile secondary feathers. |
Many juvenile (1Y) Goldfinches have 1 to 3, rarely 4 or more unmoulted old greater coverts (OGCs). These are best distinguished by the contrast in appearance of the black part of the feathers rather than the colour of the tips as illustrated in Svensson. The new (adult type) feathers have a glossy black appearance in contrast to the matt black of the juvenile feathers with the differences being a bit more obvious in males. Birds with retained old greater coverts are easy to age.
Photo 5. 1Y (Juvenile) with 2 OGCs |
Photo 6. Another juvenile with 2 OGCs |
Photo 7. 1Y (Juvenile) with 3 OCCs. |
However, some juveniles moult all their greater coverts and I found that just under 35% of the juveniles I caught had replaced them all (sample size 94). This is a significant proportion and these birds can be more difficult to age. Some ageing guides suggest there is a contrast between the new greater coverts and the old primary coverts and alula in these birds and that (with practise) this can be used for ageing. I used to agree with that view but have found a similar level of contrast between these feathers in all the known adults and therefore it isn’t a feature that stands up to close scrutiny. Having said that it is worth checking the carpal covert as this feather frequently has a dirty buff or brown fringe at the tip in juveniles although a few may be all black like most adults. In adults this feather is generally all black but very rarely a few may have a slight greyish white fringe (never dirty buff or brown as in juveniles). This feature is more obvious when the feathers are relatively fresh, in autumn and the first half of the winter, but can last through to the spring in well marked individuals.
Photo 9. A tighter crop of photo 8 above. Note there is no more contrast between the greater coverts, primary coverts and alula in this juvenile than in the adult in photo 10 below. |
The tail shape of juveniles is generally described as being more pointed than those of adults and that they usually show more wear. However, things are not quite that simple as you will see from the photos below. A large proportion of juveniles replace the central pair of tail feathers and some replace the outermost pair with the difference in wear and contrast between the black areas of the feathers identifying these birds (again the newer glossy black feathers contrasting with the more matt black juvenile tail feathers). A few may replace more tail feathers but it is very rare for juveniles to replace all their tail feathers (1 in 300 or less). I only caught 1 juvenile that had replaced all its tail and this bird was aged from the partial moult of its primaries. As ever with Goldfinches there is a lot of variation and more overlap in tail shape than is often appreciated or illustrated in ageing guides.
Photo 14. Here is another example of a 1Y (juv) tail. This bird had replaced the central and outermost pairs of tail feathers. It had also replaced all its greater coverts and the innermost tertial. |
The photos of the juvenile tails above are a fairly random selection and show the range of tail shapes that juveniles can display. Having an understanding of this variation is particularly important when determining the age of birds with no old greater coverts. All the photos were taken over a relatively short period in December and after the post juvenile moult is complete so it isn't a case of variation over a long period of time or before and after the post juvenile moult period. The level of wear and any contrast between new and old feathers are by far the most important things to look for when using the tail to age birds with shape being more of a secondary consideration.
So now for some photos of the tails of some known adults for comparison. All the photos below are of birds that were originally ringed in previous years and are not just birds that appear to show adult features.
There is a footnote in Svensson that says birds with sub-terminal white spots on the outer 3 tail feathers are adult although most adults only have spots on the outer 2. This has been repeated elsewhere but is not correct. I have caught numerous juveniles with sub-terminal white spots on the outer 3 tail feathers over the years as shown below. However, there is a paper that suggests birds with 3 white sub-terminal patches can be confidently sexed as males. That will have to be discussed in a future post on sexing Goldfinches or why you shouldn't sex most Goldfinches as my findings currently seem to suggest.
The one ageing feature that I haven't illustrated with photographs so far is the difference in primary wear that is shown by juveniles compared to adults. The primaries are always worth checking and can be very useful for confirming a birds age. In juveniles the white tips wear quite quickly in autumn whereas adults can remain relatively fresh throughout the winter.
Even though I have taken a lot of photographs it is really difficult to show some of the features well as they are quite subtle. The contrast between some feathers is fairly easy to see in good light when you can keep tilting the bird to catch the light but is much more difficult to photograph. I just hope some of you find this of interest and useful as a reference.
UPDATE 01/02/14
I caught a few Goldfinches on 30/01/14 that included another known adult (ringed in 2012) along with an unringed adult and some first winter birds. This allowed me to make a direct comparisons of the relative wear of the known adult, new adult and juveniles and to see if the differences in wear were still as marked as they had been. I took the usual range of photographs and present a few reference images below.The difference in wear is slightly greater to that seen in the late autumn/early winter in these individuals with the juvenile having lost more of its white primary tips to the point where they have almost disappeared on some feathers. The adult primary tips remain complete and very fresh in comparison and this was equally true for the new adult.
D130399 was also interesting in that I hadn't caught it in the usual way. This bird flew into a window and stunned itself on 03/11/12. I kept it in a bird bag for a short time afterwards and it seemed to recover quite quickly so I ringed it prior to release. It is nice to know that birds can fully recover from such impacts.
I will update this post again later in the year (ideally March or April) if I catch any suitable retraps/known age birds that show how the plumage wear progresses.
Photo 19. Known adult Goldfinch tail. Similar to photo 18 above. |
Photo 20. Known adult Goldfinch tail. |
Photo 21. Known adult Goldfinch tail. Again similar to photos 18, 19 and 20. This is probably the most common adult tail shape. The one thing all the adults had in common was the lack of wear to the tail feathers and all the tail feathers were clearly of the same generation.
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Photo 22. Example of a 1Y (juv) Goldfinch with white sub-terminal spots on the outer 3 tail feathers taken at Spurn in 2006. |
Photo 25. 1Y (juv) primaries showing how the white tips become quite worn and chipped by early autumn. Sometimes the white tips wear away almost completely. |
Even though I have taken a lot of photographs it is really difficult to show some of the features well as they are quite subtle. The contrast between some feathers is fairly easy to see in good light when you can keep tilting the bird to catch the light but is much more difficult to photograph. I just hope some of you find this of interest and useful as a reference.
UPDATE 01/02/14
I caught a few Goldfinches on 30/01/14 that included another known adult (ringed in 2012) along with an unringed adult and some first winter birds. This allowed me to make a direct comparisons of the relative wear of the known adult, new adult and juveniles and to see if the differences in wear were still as marked as they had been. I took the usual range of photographs and present a few reference images below.The difference in wear is slightly greater to that seen in the late autumn/early winter in these individuals with the juvenile having lost more of its white primary tips to the point where they have almost disappeared on some feathers. The adult primary tips remain complete and very fresh in comparison and this was equally true for the new adult.
Photo 31. Known adult tail. All the feathers are clearly of the same age with little or no signs of wear. |
I will update this post again later in the year (ideally March or April) if I catch any suitable retraps/known age birds that show how the plumage wear progresses.
great article, found it very informative.
ReplyDeletecheers Mike